CV


CV HIGHLIGHTS:

 Education
  • 2015  Ph.D., Physical Anthropology, New York University
  • 2008 M.A., Physical Anthropology, New York University
  • 2005 B.A., Anthropology, Summa Cum Laude, The Ohio State University
  • 2005 B.A., English, Summa Cum Laude, The Ohio State University


Peer-reviewed Publications
  • 2026. Villamil, C.I., Rodríguez, J.N., Middleton, E.R. Temporal changes in vertebral morphology of the free-ranging Rhesus macaques from Cayo Santiago, Puerto Rico. Am. J. Biol. Anthropol. 189, e70249. doi:10.1002/ajpa.70249
  • 2025. Villamil, C.I., Middleton, E.R. Modularity and integration of the neural arch and vertebral centrum in primates. Anat. Rec. 309, 105-129. doi:10.1002/ar.25653
  • 2025. Middleton, E.R., Alwell, M.T., Ward, C.V. Manubriosternal variation in anthropoid primates. Am. J. Biol. Anthropol. 186, e25053. http://dx.doi.org/10.1002/ajpa.25053
  • 2024. Villamil, C.I., Rodríguez, J.N., Middleton, E.R. Heritability in the Rhesus macaque (Macaca mulatta) vertebral column. Am. J. Primatol. e23686. doi:10.1002/ajp.23686 
  • 2024. Villamil, C.I., Middleton, E.R. Conserved patterns and locomotor-related evolutionary constraints in the hominoid vertebral column. J. Hum. Evol. 190, 103528.
  • 2023. Antón, S.C., Middleton, E.R. Making meaning from fragmentary fossils: early Homo in the Early to early Middle Pleistocene. J. Hum. Evol. 179, 103307.  doi:10.1016/j.jhevol.2022.103307
  • 2018. Ward, C.V., Maddux, S.D., Middleton, E.R. Three-dimensional anatomy of the anthropoid bony pelvis. Am. J. Phys. Anthropol. 166, 3-25.
  • 2017. Middleton, E.R., Winkler, Z.J., Hammond, A.S., Plavcan, M.J., Ward, C.V. Determinants of iliac blade orientation in anthropoid primates. Anat. Rec. 300, 810-827.
  • 2016. Antón, S.C., Taboada, H., Middleton, E.R., Rainwater, C.W., Taylor, A.B., Turner, T.R., Turnquist, J.E., Weinstein, K.J., Williams, S.A., Morphological variation in Homo erectus and the origins of developmental plasticity. Philos. Trans. R. Soc. B. 371:20150236. 
  • 2016. Williams, S.W., Middleton, E.R., Villamil, C.I, Shattuck, M.R., Vertebral numbers and human evolution. Am. J. Phys. Anthropol. 159(S61), S19-S36.


AAPA 2014 Poster presentation
Selected Abstracts and Conference Presentations
  • 2026. Middleton, E.R., Villamil, C.I. Sexual dimorphism in thoracolumbar vertebral length: patterns and insights from the hominoids. Am. J. Biol. Anthropol. 189 (S81), 128.
  • 2025. Middleton, E.R. Phenotypic integration of the thoracolumbar vertebrae and pelvis in hominoids. Am. J. Biol. Anthropol. 186 (S79), 112.
  • 2024. Villamil, C.I., Rodríguez, J.N., Middleton, E.R. Investigating size effects on heritability: a case study using the primate vertebral column. Am. J. Biol. Anthropol. 183 (S77), 187-188.
  • 2023. Middleton, E.R., Antón, S.C. Fossil pelves attributed to early Homo: variants and variation. Am. J. Biol. Anthropol. 180 (S75), 118.
  • 2017. Middleton, E.R. The modular nature of the trunk skeleton in anthropoid primates. Fed. Am. Soc. Exp. Biol. J. 31, 396.2
  • 2017. Middleton, E.R., Ward, C.V. Functional inferences from vertebral morphology and torso shape in anthropoids. Am. J. Phys. Anthropol. 162 (S64), 287.
  • 2016. Ward, C.V., Middleton, E.R. Determinants of body shape and co-variation among elements of the bony torso in anthropoids (Primates: Anthropoidea). Anat. Rec. 299, 183.
  • 2016. Ward, C.V., Middleton, E.R. Body shape and hominin origins. Am. J. Phys. Anthropol. 159 (S62), 328–329.
  • 2014. Middleton, E.R. Phenotypic patterns of rib and pelvis covariation in Old vs. New World human populations. Am. J. Phys. Anthropol. 153 (S58), 185.
Conference Symposia
  • 2017 Organizer and chair. Linked Modules in Anatomical Inference: New Approaches to Morphological Integration. Invited podium symposium, Experimental Biology annual meeting.
  • 2014 Co-organizer and chair. The Bones and Behavior Project: Fostering Integration across Biological Anthropology. Invited poster symposium, AAPA annual meeting. 
AAPA 2014 Bones and Behavior Project members - Invited Poster Symposium

Grants
  • 2016 American Association of Anatomists Postdoctoral Fellowship ($20,000)
  • 2013 Wenner-Gren Foundation Dissertation Fieldwork Grant ($15, 215)
  • 2013 Leakey Foundation Research Grant ($8,505)
  • 2013 New York University Global Research Institute Fellowship ($2,800) 
  • 2011 Margaret and Herman Sokol Predissertation Research Award ($4,000) 
  • 2009 NYCEP IGERT Summer Research and Travel Grant ($5,000)